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On OS X 10.9.5, the Calendar.app toolbar sometimes adds this button:

mystery box

There's no tooltip, and the Calendar toolbar isn't editable so there's no way to show a label for it. I can't find anything in the online documentation about it.

I think it has something to do with remote/subscribed calendars, but what, exactly?

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This is the Notifications button.

The Notifications button shows how many invitations you have. If you don’t have any invitations, it isn’t available.

Source: Calendar: Reply to an event or calendar invitation

The notification button also shows when changes are made to a subscribed calendar. In this case, the standard Accept/Decline/Maybe buttons are replaced by a single OK option to dismiss.

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