I have both Safari and Mail installed - ever since a fresh OSX Lion install. Also, I have never removed or reinstalled them or messed around with them. So this "error" message is quite a surprise to me.

How do I fix this?
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You should use the Mail.app
If this doesn't work:
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open -a Mailwork? – lhf Jul 3 '12 at 12:21Mail.appopens. – gentmatt Jul 3 '12 at 13:14